ProfessorDept. of Earth Sciences,University of California, Riverside, CA. 92521.
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Education |
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| University of Durham, U.K., Geology, 1982-1985 | |
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Visva-Bharati University, Bengali, 1985-1986 Santiniketan, West Bengal, India. |
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| University of Bristol, U.K., Geology, 1986-1990 | |
Appointments |
| Visiting Lecturer [Assistant Professor] in Paleontology, Trinity College, University of Dublin, Ireland. 1989-1990. |
| NERC (NATO) Postdoctoral Research Fellowship, Queensland Museum, Brisbane, Australia. 1990-1992. |
| Smithsonian Postdoctoral Fellow, Washington, D.C. U.S.A. 1992-1993. |
| Assistant Curator, Cincinnati Museum Center, U.S.A. 1993-1997. |
| Adjunct Assistant Professor, University of Cincinnati, U.S.A. December 1993-1997. |
| Associate Curator, Cincinnati Museum Center, U.S.A. 1997. |
| Research Associate, Department of Paleobiology, Smithsonian Institution. U.S.A. 1994-present. |
| Associate Professor, University of California, Riverside. 1997-2003. |
| Professor, University of California, Riverside. 2003-. |
Refereed Publications: |
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Hughes N.C. and Rushton A.W.A. 1990 Computer-aided reconstruction of a late Cambrian ceratopygid trilobite from Wales and its phylogenetic implications. Paleontology 33:429-445. |
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Jell P.A., Hughes N.C. and Brown A.V. 1991 Late Cambrian (post-Idamean) trilobites from the Higgins Creek area, western Tasmania. Memoirs of the Queensland Museum 30:455-485. |
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Hughes N.C. 1991 Morphological plasticity and genetic flexibility in a Cambrian trilobite. Geology 19:913-916. |
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Hughes N.C. and Droser M.L. 1992 Trace fossils from the Phe Formation (Lower Cambrian), Zanskar valley, northwestern India. Memoirs of the Queensland Museum 32:139-144. |
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Hughes N.C. and Jell P.A. 1992 A statistical/computer graphic technique for assessing variation in tectonically deformed fossils and its application to Cambrian trilobites from Kashmir. Lethaia 25:317-330. |
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Hughes N.C. 1993 Distribution, taphonomy, and functional morphology of the Upper Cambrian trilobite Dikelocephalus. Milwaukee Public Museum Contributions in Biology and Geology 84:1-49. |
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Labandeira C.C. and Hughes N.C. 1994 Biometry of the Late Cambrian trilobite Dikelocephalus, and its implications for trilobite systematics. Journal of Paleontology 68:492-517. |
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Hughes N.C. 1994 Ontogeny, intraspecific variation, and systematics of the Late Cambrian trilobite Dikelocephalus. Smithsonian Contributions to Paleobiology 79:1-89. |
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Droser M.L., Hughes N.C. and Jell P.A. 1994 Palaeoecology of Cambro-Ordovician nearshore sandstones: trace fossil evidence from Mootwingee, New South Wales, Australia. Lethaia 27:273-283. |
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Hughes N.C. 1995 Trilobite taphonomy and taxonomy: a problem and some implications. Palaios 10:283-285. |
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Hughes N.C. and Labandeira, C.C. 1995 The stability of species in taxonomy. Paleobiology 21:401-403. |
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Hughes N.C. and Chapman R.E. 1995 Growth and variation in the Silurian trilobite Aulacopleura konincki and its implications for trilobite palaeobiology. Lethaia 28:333-353. |
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Rushton A.W.A and Hughes N.C. 1996 Biometry, systematics, and biogeography of the late Cambrian trilobite Maladioidella abdita. Transactions of the Royal Society of Edinburgh: Earth Sciences 86:247-256. |
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Hughes N.C., Gunderson G.O. and Weedon M.J. 1997 Circumocular suture and visual surface of "Cedaria" woosteri (Trilobita, Late Cambrian) from the Eau Claire Formation, Wisconsin. Journal of Paleontology 71:103-107. |
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Hughes N.C. and Hesselbo S.P. 1997 Stratigraphy and sedimentology of the St. Lawrence Formation, Upper Cambrian of the northern Mississippi Valley. Milwaukee Public Museum Contributions in Biology and Geology 91:1-50. |
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Jell P.A. and Hughes N.C. 1997 Himalayan Cambrian trilobites. Special Papers in Palaeontology 58:1-113. |
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Hughes N.C. 1997 Cambrian trilobite biostratigraphy for the Himalaya. Current Science 73:736-737. |
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Fortey R.A. and Hughes N.C. 1998 Brood pouches in trilobites. Journal of Paleontology 72:638-649. |
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Webster M. and Hughes N.C. 1999 Compaction-related deformation in well-preserved Cambrian olenelloid trilobites and its implications for fossil morphometry. Journal of Paleontology 73:355-371. |
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Hughes N.C. and Cooper D.L. 1999 Paleobiologic and taphonomic aspects of the "granulosa" trilobite cluster, Kope Formation (Upper Ordovician, Cincinnati region). Journal of Paleontology 73:306-319. |
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Hughes N.C. and Jell P.A. 1999 The biostratigraphy and biogeography of Himalayan Cambrian trilobites. Geological Society of America Special Papers 328:109-116. |
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Hughes N.C. 1999 Statistical and imaging methods applied to deformed fossils. in Harper, D.A.T. (ed.). Numerial Paleobiology: computer-based modelling and analysis of fossils and their distributions. John Wiley Press, London. pp.127-155. |
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Hughes N.C. Chapman R.E. and Adrain J.M 1999 The stability of thoracic segmentation in trilobites: a case study in developmental and ecological constraints Evolution and Development 1:24-35. |
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Hughes N.C. 2000 The ecological evolution of Cambrian trilobites. in Zhuravlev A.Yu. and Riding R. (eds.). Ecology of the Cambrian Radiation. Columbia University Press, New York. p. 370-403. |
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Collier F.J. Hughes N.C. Klussendorf J. Lipps J.H. Taylor W.L. and White R.D. 2000 Fossil invertebrate and micro-fossil collections: Kinds, uses and users. Paleontological Society Special Publications 10: 25-35. 18 ms. pages. http://www.englib.cornell.edu/pri/ProBltn/fProBltn.html |
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Webster, M., Sheets, H.D., and Hughes N.C. 2001 Allometric patterning in trilobite ontogeny: Testing for allometric heterochrony in Nephrolenellus. pp 105-144. in Zelditch, M.L. ed. Beyond Heterochrony. Wiley-Liss, New York. 371 pp. |
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Hughes N.C. and Chapman R.E. 2001 Morphometry and phylogeny in the resolution of paleobiological problems Ð unlocking the evolutionary significance of an assemblage of Silurian trilobites. p. 29-54. in Adrain, J.M., Edgecombe, G.E. and Lieberman, B.S. eds. Fossils, Phylogeny, and Form: An Analytical Approach. Kluwer Academic/Plenum Press. New York. 402 pp. |
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Hughes N.C., Peng Shanchi and Luo Huilin 2002 Kunmingaspis (Trilobita) putatively from the Yunling Collage, and the Cambrian faunal history of the eastern Himalayan syntaxial region. Journal of Paleontology 76: 709-717. |
Hughes N.C. 2002 Late Middle Cambrian trace fossils from the Lejopyge armata horizon, Zanskar Valley, India, and the use of Precambrian/Cambrian ichnostratigraphy in the Indian subcontinent. Special Papers in Palaeontology 67:135-151. |
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Hughes N.C. 2003 Trilobite body patterning and the evolution of arthropod tagmosis. BioEssays. 25:386-395. |
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Minelli A., Fusco G. and Hughes N.C. 2003 Tagmata and segment specification in trilobites. Special Papers in Palaeontology. 70:31-43. |
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Peng Shanchi, Babcock L.E., Hughes N.C. and Lin Huanling. 2003 Cambrian shumardioids from northwestern Hunan, China. Special Papers in Palaeontology. 70:197-212. |
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Hughes N.C. 2003 Trilobite tagmosis and body patterning from morphological and developmental perspectives. Integrative and Comparative Biology (American Zoologist). 43:185-206. |
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Myrow P.M., Hughes N.C., Paulsen T.S., Williams I.S., Parcha S.K., Thompson K.R., Bowring S.A., Peng S.-C., and Ahluwalia A.D. 2003 Integrated tectonostratigraphic reconstruction of the Himalaya and implications for its tectonic reconstruction. Earth and Planetary Science Letters. 212:433-441. |
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Fusco G., Hughes N.C., Webster, M., and Minelli, A 2004 Exploring developmental mode in a fossil arthropo: growth and trunk segmentation in the trilobite Aulaceopleura konincki. American Naturalist. 163:167-183. |
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Hughes N.C. and Heim, N.A. 2005 Cambrian: chronology, environments, and evolution. in Selley, R, Cocks, L.R.M. and Plimer, I.R. Encylopedia of Geology. Oxford. vol. 4. pp 163-174. |
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Hughes N.C., Peng Shanchi, Bhargava, O.N., Ahulwalia, A.D., Walia, S., Myrow, P.M. and Parcha, S.K. 2005 Early Tsanglangpuan (late early Cambrian) trilobites from the Nigali Dhar syncline and the Cambrian biostratigraphy of the Tal Group, Lesser Himalaya, India. Geological Magazine, 142:57-80. |
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Jacobs, D.K., Hughes N.C., Fitz-Gibbon, S.T, and Winchell, C.J. 2005 Terminal addition, the Cambrian radiation, and the Phanerozoic evolution of bilaterian form. Evolution and Development. 7:498-514. |
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Simpson, A., Hughes N.C., Kopaska-Merkel, D.C. and Ludvigsen, R. 2005 Development of the caudal exoskeleton of the pliomerid trilobite Hintzeia plicamarginis new species. Evolution and Development 7:528-541. |
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Hughes N.C. 2005 Trilobite construction: building a bridge across the micro-macroevolutionary divide. p. 139-158. in Briggs, D.E.G. (ed.). Evolving Form and Function: Fossils and Development: Proceedings of a symposium honoring Adolph Seilacher for his contributions to paleontology in celebration of his 80th birthday. Peabody Museum of Natural History. Yale University. New Haven. |
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Myrow, P.M., Thomson, K.R., Hughes N.C., Paulsen, T.S., Sell, B.K. and Parcha, S.K. 2006 Cambrian stratigraphy and depositional history of the northern Indian margin, Spiti Valley, north-central India. Geological Society of America Bulletin. 118:491-510. |
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Myrow, P.M., Snell, K.E., Hughes N.C., Paulsen, T.S., Heim, N.A. and Parcha, S.K. 2006 Cambrian depositional history of the Zanskar Valley region of the Indian Himalaya: tectonic implications. Journal of Sedimentary Research 76:364-381. |
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Hunda, B.R., Hughes N.C., and Flessa, K. 2006 Trilobite taphonomy and temporal resolution in the Mt. Orab Shale (Upper Ordovician, Ohio, U.S.A.) Palaios 21:26-45. |
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Hughes N.C., Minelli A., and Fusco G. 2006 The ontogeny of trilobite segmentation: a comparative approach. Paleobiology 32:603-628. |
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Paulsen, T.S., Demosthenous, C.M., Myrow, P.M., Hughes, N.C., and Parcha, S.K. 2007 Paleostrain stratigraphic analysis of calcite twins across the Cambrian-Ordovician unconformity in the Tethyan Himalaya, Spiti and Zanskar valley regions, India. Journal of Asian Earth Sciences 31:44-54. |
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Hughes, N.C. 2007 The Evolution of Trilobite Body Patterning. Annual Reviews in Earth and Planetary Sciences 35:401-434. |
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Hunda B.R and Hughes N.C. 2007 Evaluating paedomorphic heterochrony in trilobites: the case of the diminutive trilobite Flexicalymene retrorsa minuens from the Cincinnatian Series (Upper Ordovician), Cincinnati region. Evolution and Development 9:483-496. |
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Zhu X.-J., Hughes N.C., and Peng S.-C. 2007 On a new species of Shergoldia Zhang & Jell 1987 (Trilobita), the family Tsinaniidae, and the Order Asaphida. Memoirs of the Australasian Association of Palaeontologists 34:243-253. |
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Hughes, N.C. 2007 Strength in numbers: high phenotypic variance in early Cambrian trilobites and its evolutionary implications. BioEssays 29:1081-1084. |
Hughes N.C., Haug J.T. and Waloszek D. 2008 Basal euarthropod development: a fossil-based perspective. p. 279-296. in Minelli A. and Fusco G. eds. Evolutionary Pathways Ð Key Themes in Evolutionary Developmental Biology. Cambridge University Press. 426pp. |
COPIES OF RECENT PAPERS CAN BE DOWNLOADED FROM http://earthsciences.ucr.edu/hughes.html |
Editorially Reviewed Papers: |
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Hughes N.C. and Fortey R.A. 1995 Sexual dimorphism in trilobites, with an Ordovician case study, 419-421. in Cooper J.C., Droser M.L. and Finney S.C. eds. Ordovician Odyssey, SEPM Pacific Section, Los Angeles, CA. 499p. |
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Droser M.L. and Hughes N.C. 1995 Infaunal communities and tiering in Ordovician shallow marine terrigenous clastic and carbonate settings: Ichnofabric and trace fossil evidence, 405-408. in Cooper J.C., Droser M.L. and Finney S.C. eds. Ordovician Odyssey, SEPM Pacific Section, Los Angeles, CA. 499p. |
Refereed Comments: |
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Hughes N.C and Hughes S.M. 1993 Development of the zootype. Nature 363:307. |
Professional Service: |
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| Member of Joint Technical Program Committee of GSA | |
| Member of Paleontological Society's Committee on Collections | |
| Coordinator of Paleontological Society's Fossil Festival, Cincinnati | |
| Co P.I. and organizer of national NSF workshop on curatorial standards and databases in invertebrate paleontology, Washington D.C. | |
| Panel member. Biotic Surveys and Inventories program. National Science Foundation. | |
| Panel member. Research Collections in Systematics and Ecology program. National Science Foundation. | |
| Paleontological Society's Distinguished Speaker Program. | |
| Member, Student Grants Committee, Paleontological Society | |
| Technical Editor, Journal of Paleontology. | |
| Councilor-at-large, under 40, Paleontological Society | |